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Abstract
Vertical growth is an important element to consider when evaluating the movement of an alpine treeline. The vertical growth of trees is decisive in the establishment of trees upslope of the existing treeline, as trees must be able to grow up, mature, and reproduce in order for the treeline to advance. The purpose of this study was to explore the possible causes of, and factors influencing, the vertical growth of trees in a treeline environment, specifically at the alpine treeline of Pike's Peak, CO. Vertical growth was first studied on an individual scale, specifically investigating the thermal regime of trees and its impact on growth. The air temperature profile showed a nighttime inversion of daytime conditions. During the night there was a lapse rate of approximately 1°C, with the coldest conditions closest to the ground. Thus, the smallest trees were in significantly colder environments during the night than the largest trees. During the day, there was a lapse rate of approximately 3°C per meter, a very high lapse rate, with the warmest conditions occurring closest to the ground. Thus, the smallest trees were in the warmest conditions throughout the day. Additionally, it was found that small trees were coupled to ground conditions during the day as well as the night, and that the taller trees were coupled to atmospheric conditions. Yet, the coupling relationships were not exact, as the tree temperatures never exactly matched the ground or atmospheric temperatures. Finally, I investigated whether daytime or nighttime temperatures impacted growth more closely. It was found that daytime conditions were more important for the growth of trees at the study site on Pike's Peak. The second part of the study investigated tree growth on a stand-wide scale, considering whether or not there were larger spatial patterns affecting the vertical growth of trees. I found that a shelterbelt-like system was in place at the treeline, the presence of which seemed to be affecting the growth of the trees within its bounds. Specifically, there was depression of growth directly upslope of the trees creating the upper bounds of the treeline, then an area of facilitated growth, ending with a return to normal conditions. Yet, these shelterbelt conditions were only detected for trees one meter or taller. The growth patterns for trees under 1 meter did not correlate to the growth patterns of taller trees. Additionally, the shelterbelt conditions would only be present during the day, which further confirms the importance of daytime conditions found in the first study. This exploratory study was a first look into the drivers of vertical growth of trees at an alpine treeline.